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Josh is initially upset with Lou for hiring Donna, and he implies in a tense conversation with Donna that he is reluctant to hire her since he feels betrayed by her decision to work for a competing candidate which he, her "mentor", did not support.
The two reconcile on the campaign trail, with their relationship becoming romantic during the season 7 episode "The Cold".
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Although all the senior staff's assistants are continuing characters with personal backgrounds, Donna is the best defined and most often featured staff member on the assistant level, and her difficult, semi-romantic relationship with Josh is a major plot device during later seasons.
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However, her citizenship is quickly restored after the discovery of a grandfather clause in the law. Donna moved to Wisconsin, where she attended University of Wisconsin–Madison and dropped out halfway through to support her boyfriend as he went through medical school. She then left Madison and drove to New Hampshire to work for the "Bartlet for America" campaign, even going so far as to pretend she'd already been hired as Josh Lyman's assistant.
Josh discovers the deception but is somewhat amused by it and, impressed by her initiative, hires her.